List of human evolution fossils
The following tables give an overview of notable finds of hominin fossils and remains relating to human evolution , beginning with the formation of the tribe Hominini (the divergence of the human and chimpanzee lineages ) in the late Miocene , roughly 7 to 8 million years ago.
As there are thousands of fossils, mostly fragmentary, often consisting of single bones or isolated teeth with complete skulls and skeletons rare, this overview is not complete, but shows some of the most important findings. The fossils are arranged by approximate age as determined by radiometric dating and/or incremental dating and the species name represents current consensus; if there is no clear scientific consensus the other possible classifications are indicated.
The early fossils shown are not considered ancestors to Homo sapiens but are closely related to ancestors and are therefore important to the study of the lineage. After 1.5 million years ago (extinction of Paranthropus ), all fossils shown are human (genus Homo ). After 11,500 years ago (11.5 ka, beginning of the Holocene ), all fossils shown are Homo sapiens (anatomically modern humans ), illustrating recent divergence in the formation of modern human sub-populations .
The chimpanzee–human divergence likely took place around 10 to 7 million years ago.[ 1] The list of fossils begins with Sahelanthropus , dated some 7 million years ago, which may or may not be ancestral to both the human and the chimpanzee lineage, although its status as a hominin or a hominin ancestor remain debated. For the earlier history of the human lineage, see Timeline of human evolution#Hominidae , Hominidae#Phylogeny .
The Trachilos footprints from the island of Crete as well as the Graecopithecus fossils are not included in the list due to taxonomic uncertainty.
Image
Name
Age (Ma )
Species
Year discovered
Country
Discovered by
Now located at
Ardi
4.40[ 5]
Ardipithecus ramidus
1994
Ethiopia
Yohannes Haile-Selassie
Lothagam mandible (KNM-LT 329)[ 6]
4.60±0.40[ 7]
Australopithecus sp. or undetermined Hominidae
1967
Kenya
Arnold Lewis,[ 8] Bryan Patterson [ 9] [ 10] [ 11]
KNM-TH 13150
4.70±0.55[ 12]
Orrorin praegens
1984
Kenya
Kiptalam Cheboi[ 9]
KNM-KP 271
4.00[ 13]
Australopithecus anamensis
1965
Kanapoi, Kenya
Bryan Patterson[ 9]
Laetoli Footprints
3.70
Bipedal hominin
1976
Tanzania
Mary Leakey
LH 4
3.40±0.50
Australopithecus afarensis
1974
Laetoli , Tanzania
Mary Leakey [ 14]
KSD-VP-1/1 (Kadanuumuu )
3.58
Australopithecus afarensis
2005
Ethiopia
Yohannes Haile-Selassie
KT-12/H1 (Abel)
3.50
Australopithecus bahrelghazali
1995
Chad
Mamelbaye Tomalta and Michel Brunet
N'Djamena (Chad), BEAC
KNM-WT 22944 G-J [ 15]
3.50
Australopithecus sp.
1990
Kenya
Multinational team
National Museums of Kenya
KNM-WT 40000 (Flat Faced Man) [ 16]
3.50-3.20
Kenyanthropus platyops
1999
Lake Turkana (West Lake Turkana), Kenya
Justus Erus and Meave Leakey [ 17]
BRT-VP-3/14
3.40±0.10
Australopithecus deyiremeda
2015
Ethiopia
Yohannes Haile-Selassie[ 18]
Stw 573 (Little foot)
3.67
Australopithecus prometheus (?)
1994
Sterkfontein , South Africa
Ronald J. Clarke
DIK-1 (Selam)
3.30
Australopithecus afarensis
2000
Ethiopia
Zeresenay Alemseged
AL 288-1 (Lucy)
3.20
Australopithecus afarensis
1974
Ethiopia
Tom Gray, Donald Johanson , Yves Coppens and Maurice Taieb
National Museum of Ethiopia
AL 200-1
3.10±0.10
Australopithecus afarensis
1975
Afar Region , Ethiopia
Donald Johanson Yves Coppens and Maurice Taieb
AL 129-1
3.10±0.10
Australopithecus afarensis
1973
Afar Region , Ethiopia
Donald Johanson
AL 444-2 [ 19]
3.00
Australopithecus afarensis
1992
Afar Region , Ethiopia
Yoel Rak
LD 350-1 [ 20]
2.775±0.025[ 21]
Homo (?)
2013
Ethiopia
Chalachew Seyoum
Taung Child 1
3.03–2.61
Australopithecus africanus
1924
Buxton-Norlim Limeworks, South Africa
Raymond Dart
University of the Witwatersrand
Name
Age (Ma )
Species
Date discovered
Country
Discovered by
Now located at
KNM-WT 17000 (The Black Skull)
2.50
Paranthropus aethiopicus
1985
Kenya
Alan Walker
BOU-VP-12/130[ 22]
2.50
Australopithecus garhi
1997
Ethiopia
Yohannes Haile-Selassie
LD-760[ 23]
2.63
Australopithecus sp. indet.
2025
Ethiopia
STS 71 [ 24]
2.61–2.07
Australopithecus africanus
1947
Sterkfontein , South Africa
Robert Broom and John T. Robinson
Ditsong National Museum of Natural History
STS 52
2.61–2.07
Australopithecus africanus
1947
Sterkfontein , South Africa
Robert Broom
Ditsong National Museum of Natural History
UR 501 (Uraha jawbone)
2.40±0.10
Homo rudolfensis [ 25]
1991
Malawi
Tyson Msiska, Timothy Bromage, Friedemann Schrenk
STS 5 (Mrs. Ples) (STS 14 )[ 26]
2.07[ 27]
Australopithecus africanus
1947
Sterkfontein , South Africa
Robert Broom
Ditsong National Museum of Natural History
DNH 134 (Simon)[ 28]
2.04[ 28]
Homo erectus
2015
Drimolen Main Quarry, South Africa
Andy Herries' team (excavated by Richard Curtis, Andy Herries, Angeline Leece; reconstructed by Jesse Martin)
University of the Witwatersrand
DNH 155[ 29]
2.04–1.95
Paranthropus robustus
2018
Drimolen Main Quarry, South Africa
Andy Herries and Stephanie Baker's team (first found by Samantha Good and excavated by Samantha Good, Angeline Leece, Stephanie Baker and Andy Herries; reconstructed by Jesse Martin)
University of the Witwatersrand
DNH 152[ 28] (Khethi)
2.04–1.95[ 28]
Paranthropus robustus
2018
Drimolen Main Quarry, South Africa
Andy Herries and Stephanie Baker's team (first part found by Khethi Nkosi. later parts by Amber Jaeger, Eunice Lalunio; reconstructed by Jesse Martin & Angeline Leece)
University of the Witwatersrand
DNH 7 (Eurydice) [ 30]
2.04–1.95[ 28]
Paranthropus robustus
1994
Drimolen , Drimolen Main Quarry, South Africa
R. Smith and André Keyser
University of the Witwatersrand
KNM-ER 64060
2.03
Homo habilis
2012
Ileret , Kenya
KNM-ER 64061
2.02
Homo erectus
2012-2013
Ileret , Kenya
TM 1517 [ 31]
2.0
Paranthropus robustus
1938
South Africa
Gert Terblanche
Ditsong National Museum of Natural History
MH1 (Karabo)[ 32] [ 33]
1.98[ 34]
Australopithecus sediba
2008
Malapa , South Africa
Matthew Berger and Lee Rogers Berger
University of the Witwatersrand
KNM-ER 1813
1.90
Homo habilis
1973
Kenya
Kamoya Kimeu
KNM-ER 1470
1.90
Homo rudolfensis
1972
Kenya
Bernard Ngeneo [ 35]
SK 48
2.25–1.80
Paranthropus robustus
1948
Swartkrans , South Africa
Robert Broom
Ditsong National Museum of Natural History
SK 46 [ 36]
2.25–1.80
Paranthropus robustus
1949
Swartkrans , South Africa
Robert Broom
Ditsong National Museum of Natural History
SK 847 [ 37]
2.25–1.80
Homo habilis
1949
Swartkrans , South Africa
Ditsong National Museum of Natural History
OH 24 (Twiggy) [ 38]
1.80
Homo habilis
1968
Tanzania
Peter Nzube
OH 8 [ 39]
1.80
Homo habilis
1960
Olduvai , Tanzania
D2700 (Dmanisi Skull 3 )
1.81±0.40[ 40]
Homo erectus
2001
Dmanisi , Georgia
David Lordkipanidze and Abesalom Vekua
D3444 (Dmanisi Skull 4 )
1.81±0.40
Homo erectus
2003
Dmanisi , Georgia
David Lordkipanidze
D4500 (Dmanisi Skull 5 )
1.81±0.40
Homo erectus
2005 (published in 2013)
Dmanisi , Georgia
David Lordkipanidze
KNM-ER 62000–62003[ 41]
1.84±0.60
Homo rudolfensis
2012
Koobi Fora , Kenya
Meave Leakey 's team
KNM-ER 64062
1.84±0.02
Homo erectus
2013
Ileret , Kenya
OH 5 (Zinj or nutcracker man)
1.75
Paranthropus boisei
1959
Tanzania
Mary Leakey
OH 7 (Johnny's Child)
1.75
Homo habilis
1960
Tanzania
Jonathan and Mary Leakey
StW 53
1.8–1.6[ 27]
variously A. africanus , H. habilis , H. gautengensis
1976
Sterkfontein , South Africa
A. R. Hughes
University of the Witwatersrand
KNM-ER 1805
1.74
Homo habilis
1973/4
Kenya
Paul Abell
Yuanmou Man
1.70 or 0.60–0.50 (disputed)[ 42]
Homo erectus
1965
China
Fang Qian
KNM-ER 406
1.70
Paranthropus boisei
1969
Kenya
Richard Leakey
KNM-ER 732[ 43]
1.70
Paranthropus boisei
1970
Kenya
Richard Leakey
KNM-ER 23000[ 44]
1.70
Paranthropus boisei
1990
Koobi Fora , Kenya
Benson Kyongo
KNM-WT 17400[ 45] [ 46]
1.70
Paranthropus boisei
Not known[ 47]
Lake Turkana (West Lake Turkana) Kenya
unknown[ 47]
National Museums of Kenya, Nairobi (Kenia)
KNM-ER 3733
1.63±0.15[ 48]
Homo ergaster (a.k.a. African Homo erectus )
1975
Kenya
Lantian Man
1.62±0.03
Homo erectus
1963
Lantian County , China
Woo Ju-Kang
KNM-WT 15000 (Turkana Boy)
1.60
Homo ergaster (a.k.a. African Homo erectus )
1984
Lake Turkana (West Lake Turkana), Kenya
Kamoya Kimeu
Kenya National Museum
Peninj Mandible
1.50
Paranthropus boisei
1964
Tanzania
Richard Leakey
Ileret Footprints
1.50
Homo erectus
2007-2014
Ileret , Kenya
KNM-ER 992
1.50
Homo ergaster (a.k.a. African Homo erectus )
1971
Kenya
Richard Leakey
KNM-ER 3883
1.57±0.08
Homo erectus
1976
Kenya
Richard Leakey
Mojokerto 1 (Mojokerto child)
1.43±0.10
Homo erectus
1936
Indonesia
Andojo, G.H.R. von Koenigswald
BL02-J54-100 [ 49]
1.40
Similar to H. heidelbergensis
Unknown
Spain
Unknown
KGA 10-525 [ 50] [ 51]
1.40
Paranthropus boisei
1993
Konso-Gardula, Ethiopia
A. Amzaye
OH 9 (Chellean Man) [ 52]
1.40
Homo erectus
1960
Olduvai , Tanzania
Louis Leakey
Sima del Elephante maxilla [ 53]
1.40
Homo erectus ?
2022
Spain
OH 80
1.34
Paranthropus boisei
2010
Olduvai Gorge , Tanzania
Manuel Dominguez-Rodrigo
National Museum of Tanzania
ATE9-1 [ 54]
1.20
Homo sp. or Homo erectus ?[ 53]
2008
Spain
Eudald Carbonell
Museo de la Evolución Humana , Burgos (Spain)
Kocabaş
1.10[ 55]
Homo erectus [ 56]
2002
Turkey
M. Cihat Alçiçek
Daka
1.00
Homo erectus
1997
Ethiopia
Henry Gilbert
Sangiran 4
1.00
Homo erectus
1939
Indonesia
G.H.R. von Koenigswald
Sangiran 2
1.15±0.45
Homo erectus
1937
Indonesia
G.H.R. von Koenigswald
Madam Buya [ 57]
1.00
Homo erectus
1997
Eritrea
Ernesto Abbate
National Museum of Eritrea
ATD6-15 and ATD6-69
(Niño de la Gran Dolina 342)
0.900[ 58]
Homo antecessor orHomo erectus
1994
Spain
Bermúdez & Arsuaga
Museo de la Evolución Humana , Burgos (Spain)
Trinil 2 Pithecanthropus-1 or Java Man [ 59]
0.850±0.150
Homo erectus
1891
Indonesia
Eugène Dubois
Naturalis Biodiversity Center , Leiden
Ternifine 2-3 now Tighennif [ 60]
0.70
Homo erectus
1954
Algeria
C. Arambourg & B. Hoffstetter
Sangiran 17[ 61]
0.70
Homo erectus
1969
Indonesia
S. Sartono
Peking Man
0.73±0.50[ 62]
Homo erectus
1921
China
Davidson Black
Lost/stolen
Heidel
0.60
Homo Aeserniensis or Homo Heidelbergensis
2014
Italy
National Paleolithic Museum of Isernia
Nanjing Man
0.60±0.02
Homo erectus
1993
China
Liu Luhong
Bodo [ 63]
0.600
Homo heidelbergensis orHomo erectus
1976
Ethiopia
A. Asfaw
Benjamina [ 64] [ 65]
0.53
Homo neanderthalensis [ 66]
2001-2001
Spain
Ana Gracia Téllez
Mauer 1 (Heidelberg Man)
0.50
Homo heidelbergensis
1907
Germany
Daniel Hartmann
Institute of Earth Sciences, Heidelberg University
Saldanha man [ 67]
0.50
Homo rhodesiensis
1953
South Africa
Boxgrove Man
0.50[ 68]
Homo heidelbergensis
1994
UK
Natural History Museum
Arago 21 (Tautavel Man)
0.45
Homo erectus
1971
France
Henry de Lumley
Ceprano Man [ 69] [ 70]
0.450±0.050
Homo cepranensis /Homo heidelbergensis
1994
Ceprano , Italy
Italo Biddittu
Servizio di antropologia, Soprintendenza ai beni culturali, Regione Lazio, Italy
Agamenón [ 71]
0.43
Homo neanderthalensis [ 66]
1997
Spain
Paleontological teams
Museo de la Evolución Humana , Burgos (Spain)
Miguelón
0.40
Homo neanderthalensis [ 66]
1992
Spain
Bermúdez , Arsuaga & Carbonell
Museo de la Evolución Humana , Burgos (Spain)
Aroeira 3
0.40
Homo heidelbergensis
2014
Portugal
João Zilhão [de ]
Museu Nacional de Arqueologia, Lisbon
Salé cranium [ 72] [ 73] [ 74]
0.40-0.20
Homo sapiens ?[ 75]
1971
Morocco
Quarry worker
Swanscombe Man [ 76]
0.40
Homo neanderthalensis
1935, 1936, 1955
UK
Alvan T Marston, John J Wymer and Adrian Gibson
Natural History Museum
Ndutu [ 77] [ 78]
0.45±.04
Homo neanderthalensis affinities
1973
Tanzania
Amini Aza Mturi
Hexian Man [ 79] [ 80]
0.412±0.025[ 81]
Homo erectus
1980-1981[ 82]
Hexian , China
BH -1
0.40[ 83]
Homo heidelbergensis
Mala Balanica , Serbia
Gawis cranium
0.350±0.150
Homo erectus /Homo sapiens
2006
Ethiopia
Asahmed Humet
Steinheim Skull
0.35
Homo heidelbergensis
1933
Germany
Dinaledi Chamber hominins
0.325±0.090[ 84]
Homo naledi
2013
South Africa
Rick Hunter and Steven Tucker
University of the Witwatersrand (South Africa)
Montmaurin-La Niche mandible
~0.25
Homo heidelbergensis
1949
Haute-Garonne , France
Louis Mèroc
Musée de l'Homme
Name
Age (ka )
Species
Year discovered
Country
Discovered by
Now located at
Dragon Man
309–138
Homo longi
1933
China
Hebei GEO University
Broken Hill 1 (Kabwe 1, Rhodesian Man)
299±25[ 85]
Homo rhodesiensis (Homo heidelbergensis )
1921
Zambia
Tom Zwiglaar
Jebel Irhoud 1–5
315±32[ 86]
Homo sapiens
2017
Morocco
INSAP
Samu [ 87]
275±25
Homo heidelbergensis
1964
Hungary
László Vértes
Dali Man [ 88]
260±20[ 89]
Homo daliensis
1978
China
Shuntang Liu
Jinniushan
260-200 ka[ 90]
Homo longi Homo daliensis
1984
China [ 91]
Paleolithic Archeology Student Excavation Team[ 91]
Florisbad Skull
259±35
early Homo sapiens or Homo heidelbergensis or Homo helmei
1932
South Africa
T. F. Dreyer, G. Venter
Galilee Man
250±50
Homo heidelbergensis
1925
Israel
Francis Turville-Petre
Coupe-Gorge [ 92]
250
Homo heidelbergensis
1949
France
Raoul Cammas
Montmaurin-La Niche mandible [ 93]
250
Homo heidelbergensis
1949
France
Raoul Cammas
Musée de l'Homme
Saccopastore 1
250[ 94]
Homo neanderthalensis
1929
Grotta Guattari / Italy
Mario Grazioli
Saccopastore 2 [ 95]
250
Homo neanderthalensis
1935
Grotta Guattari / Italy
Henry Breuil and Alberto Carlo Blanc
Narmada Human
236-46
Homo erectus or Homo sapiens
1982
Narmada River , India
Arun Sonakia
Bontnewydd Palaeolithic site , Denbighshire, Wales
230
Homo neanderthalensis
1981
Wales , UK
Apidima 1 (LAO 1/S1)
210[ 96]
Homo sapiens
1978
Apidima Cave / Greece
Theodore Pitsios[ 97]
Petralona 1
200±40[ 98]
Homo heidelbergensis (uncertain)
1960
Greece
Omo remains
233±22[ 99] or 195±5[ 100]
Homo sapiens
1967
Ethiopia
Richard Leakey
Laterite Baby
190(?)[ 101]
H. erectus orH. sapiens
2001
Tamil Nadu , India
P Rajendran
Misliya-1
187±13[ 102]
Homo sapiens
2002
Israel
Israel Hershkovitz
Apidima 2 (LAO 1/S2)
170[ 96]
Homo neanderthalensis
1978
Apidima Cave / Greece
Theodore Pitsios[ 97]
Penghu 1
160±30 or 40±30[ 103]
Denisovan ?[ 104] [ 105] [ 106]
c. 2008
Taiwan
National Museum of Natural Science
Herto remains [ 107]
160
Homo sapiens
1997
Ethiopia
Tim White
Xiahe mandible
160[ 108]
Denisovan
1980
China
Altamura Man
151±21[ 109]
Homo neanderthalensis
1993
Italy
in situ
Nesher Ramla Homo
140±120
Nesher Ramla Homo orHomo neanderthalensis
2021
Israel
Israel Hershkovitz
Maba Man
140±120
early modern human ,
Homo neanderthalensis
or
Denisovan
1958
Shaogun , China
Institute of Vertebrate Paleontology and Paleoanthropology
Callao Man (CCH9)
134±14[ 110]
Homo luzonensis
2011–2015
Callao Cave , Philippines
Florent Détroit & Armand Mijares
EH 06
132–88
Homo sapiens ?
2002–2006
Lake Eyasi , Tanzania
Manuel Domínguez-Rodrigo
LH 18
120±30
Homo sapiens
1976
Ngaloba beds at Laetoli , Tanzania
Mary Leakey [ 111]
Tabun C1 [ 112]
120
Homo neanderthalensis
1967
Israel
Arthur Jelinek
Sarstedt (Sst) I-III [ 113]
115-58 ka
Homo neanderthalensis ?
1997-1999
Germany
Frangenberg brothers
Krapina 3 [ 114]
113.5±13.5[ 115]
Homo neanderthalensis
1899
Croatia
Dragutin Gorjanović-Kramberger
Ngandong 7
112[ 116]
Homo erectus
1931
Indonesia
C. ter Haar and G. H. R. von Koenigswald
Denisova 8
110[ 117] [ 118]
Denisovan
2010
Russia
Qafzeh 6 [ 119]
95±5[ 115]
Homo sapiens
1930
Israel
R. Neuville, M. Stekelis
Qafzeh 9
100–90[ 115]
Homo sapiens [ 120] [ 121]
1933
Israel
B. Vandermeersch
Scladina
103±23[ 115]
Homo neanderthalensis
1993
Belgium
Skhul 5
100±20
Homo sapiens
1933
Israel
T. McCown and H. Moivus Jr.
Skhul 9
100±20
Homo sapiens
Israel
Klasies River Caves [ 122]
100±25
Homo sapiens
1960
South Africa
Ray Inskeep, Robin Singer, John Wymer, Hilary Deacon
Eve's footprints
117
Homo sapiens
1995
South Africa
David Roberts & Lee R. Berger
Liujiang man
33-23
Homo sapiens
1958
China
Denny [ 123] [ 124] [ 125]
90
Hybrid – (Homo neanderthalensis /Denisovan )
2012
Denisova Cave / Siberia / Russia
Viviane Slon & Svante Pääbo
Max Planck Institute for Evolutionary Anthropology (Leipzig, Germany )
Panga ya Saidi [ 126]
78.3±4.1
Homo sapiens
2021
Kenya
Obi-Rakhmat 1 [ 127]
75[ 115]
Homo neanderthalensis
2003
Uzbekistan
Teshik-Tash Skull [ 128]
70
Homo neanderthalensis
1938
Uzbekistan
A. Okladnikov
La Ferrassie 1
70
Homo neanderthalensis
1909
France
R. Capitan and D. Peyrony
Shanidar 1
70±10
Homo neanderthalensis
1961
Iraq
Ralph Solecki
Sambungmacan (Sm) 1-4
70- 40[ 129]
Homo erectus
1973-2001[ 129]
Indonesia
Construction and fossil collectors[ 129]
Callao Man (CCH1)
66.7±1[ 110]
Homo luzonensis
2007
Callao Cave , Philippines
Florent Détroit & Armand Mijares
La Quina 5 [ 130]
65
Homo neanderthalensis
France
La Chapelle-aux-Saints 1
60
Homo neanderthalensis
1908
France
A. and J. Bouyssonie and L. Bardon
Kebara 2 (Moshe)[ 131]
60
Homo neanderthalensis
1983
Israel
Lynne Schepartz
Amud 7 [ 132]
55±5
Homo neanderthalensis
Israel
LB 1 (Hobbit)
55±5
Homo floresiensis
2003
Liang Bua , Indonesia
Peter Brown
Manot 1
55
Homo sapiens
2008
Israel
[ 133]
La Quina 18 [citation needed ]
52.5±7.5[ 115]
Homo neanderthalensis
France
Tam Pa Ling Cave [ 134]
54.5±8.5[ 134] [ 135]
Homo sapiens
2009
Laos
Name
Age (ka )
Species
Date discovered
Country
Discovered by
Now located at
Mungo Man
50±10
Homo sapiens
1974
Australia
Mt. Circeo 1 [ 136]
50±10
Homo neanderthalensis
1939
Italy
Prof. Blanc
SID-00B
49.2±2.5[ 137]
Homo neanderthalensis
1994
Sidrón Cave , Spain
Simanya Neanderthals [ 138]
49-42
Homo neanderthalensis
1978-1979, 2022
Simanya cave , Spain
Miguel Aznar
Archaeology Museum of Catalonia
Callao Man (CCH6a–e)
49.6±1.2[ 110]
Homo luzonensis
2011–2015
Callao Cave , Philippines
Florent Détroit & Armand Mijares
Tabon tibia (IV-2000-T-197)
47±11
Homo sapiens
2000
Tabon Caves , Philippines
National Museum of Anthropology [ 139]
Ust'-Ishim man
45
Homo sapiens
2008
Russia
Nikolai Peristov
Kents Cavern 4 maxilla
43.5±2.5
Homo sapiens
1927
UK
Zlatý kůň woman
43
Homo sapiens
1950
Czech Republic
Tianyuan man
40.5±1.5
Homo sapiens
2007
China
Amud 1 [ 140]
41[ 141]
Homo neanderthalensis
1961
Israel
Hisashi Suzuki
Neanderthal 1 [ 142]
40
Homo neanderthalensis
1856
Germany
Johann Carl Fuhlrott
Denisova hominin (X-Woman)
40
Homo sp. Altai
(also known as Homo longi )[ 143] [ 144] [ 145]
2008
Russia
Johannes Krause, et al.
hominin toe bone
40
Homo sp. Altai (possible Neanderthal –Denisovan hybrid)
2010
Russia
Oase 1
42–37[ 146]
Homo sapiens (EEMH x Neanderthal hybrid)
2002
Romania
Kostenki-14 (Markina Gora)
40–37[ 147]
Homo sapiens (EEMH )
1954
Russia
SID-20[ 148]
37.30±0.83[ 137]
Homo neanderthalensis
1994
Sidrón Cave , Spain
Balangoda Man
37
Homo sapiens
2012
Sri Lanka
Hofmeyr Skull
36
Homo sapiens
1952
South Africa
Wadjak 1 [ 149]
33±4.5[ 150]
Homo sapiens (proto-Australoid [ 151] )
1888
Indonesia
Red Lady of Paviland
33
Homo sapiens
1823
Wales , UK
William Buckland
Yamashita-Cho Man
32
Homo sapiens
1962
Japan
Engis 2
40±10[ 115] [ 152]
Homo neanderthalensis
1829
Belgium
Philippe-Charles Schmerling
Gibraltar 1
40±10[ 115]
Homo neanderthalensis
1848
Gibraltar
Captain Edmund Flint
Natural History Museum, London
Le Moustier
40±10
Homo neanderthalensis
1909
France
Denisovan tooth
40±10
Homo sp. Altai
2000
Russia
PES-2
38.9–92
Uncertain, possibly Homo neanderthalensis
Serbia
PES-1[ 153]
31–29
Uncertain, possibly Homo sapiens
Serbia
Yana RHS
31.63
Homo sapiens
Russia
Sungir I
30.25±0.25
Homo sapiens
Russia
Cro-Magnon 1
30
Homo sapiens (EEMH )
1868
France
Louis Lartet
WLH-50
29±5
Homo sapiens
1982
Australia
Pangpond [ 154]
29
Homo sapiens
2025
Khao Sam Roi Yot , Thailand[ 155]
Predmost 3 [ 156]
26
Homo sapiens
1894
Czech Republic
Karel Jaroslav Maška
Lapedo Child
24.5
Homo neanderthalensis orHomo sapiens
1998
Portugal
João Zilhão
Mid-Upper Paleolithoic human humerus from Eel Point, Caldey Island , Wales, UK[ 157]
24
Homo sapiens
1997
Wales , UK
MA-1 (Mal'ta boy)
24
Homo sapiens (ANE )
1920s
Russia
Abri Pataud Woman
20.6
Homo sapiens
France
[ 158]
Minatogawa 1
17±1
Homo sapiens
1970
Japan
Anthropology Museum, Tokyo University
Tandou [ 159] [ 160]
17
Homo sapiens
1967
Australia
Duncan Merrilees
Tabon skull cap (P-XIII-T-288)
16.5±2
Homo sapiens
1962
Tabon Caves , Philippines
Robert B. Fox
National Museum of Anthropology [ 139]
Gough's Cave [ 161] [ 162]
14.7
Homo sapiens
2010
UK
Iwo Eleru skull
13[ 163]
Homo sapiens
1965
Nigeria
"Kotias"[ 164]
13
Homo sapiens (CHG )
Kotias Klde cave , Georgia
Arlington Springs Man
13[ 165]
Homo sapiens
1959
United States
Phil Orr
Chancelade find
14.5±2.5[ 166]
Homo sapiens
1888
France
Villabruna 1
14
Homo sapiens (WHG )
1988
Italy
Bonn-Oberkassel double burial [ 167]
14-13[ 167]
Homo sapiens
1914[ 168]
Germany
Bichon man
13.7
Homo sapiens (WHG )
1956
Switzerland