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Chorrillo Formation

Chorrillo Formation
Stratigraphic range: Maastrichtian
Satellite image of the Chorrillo Formation site (in the middle on the right). In the bottom left is the Perito Moreno Glacier.
TypeGeological formation
UnderliesCalafate Formation
OverliesLa Irene Formation
Thickness>50 m (160 ft)
Lithology
PrimaryClaystone
Location
LocationPatagonia
Coordinates49°54′S 72°30′W / 49.9°S 72.5°W / -49.9; -72.5
Approximate paleocoordinates51°48′S 62°12′W / 51.8°S 62.2°W / -51.8; -62.2
RegionSanta Cruz Province
CountryArgentina
ExtentSoutheast of Viedma Lake
North of Argentino Lake
Austral Basin
Type section
Named byArbe & Hechem
Year defined1984
Chorrillo Formation is located in Argentina
Chorrillo Formation
Chorrillo Formation (Argentina)

The Chorrillo Formation, also named as Chorillo Formation,[1] is a Maastrichtian (Late Cretaceous Epoch, 72.1–66 million years ago) geologic formation in southern Patagonia, Argentina. The formation is more than 50 metres (160 ft) thick and underlies the Calafate Formation and rests on top of the La Irene Formation.[2][3] It constitutes a significant uppermost Cretaceous continental sedimentary sequence within the Magallanes Basin, located in Santa Cruz Province, Argentina, approximately 30 km southwest of El Calafate. Initial investigations in the 1990s laid the groundwork, with detailed paleontological studies emerging since 2019, revealing a rich biotic assemblage.[4] The formation correlates with the lower Dorotea Formation in Chile, sharing dinosaurian ecotypes, while both are clearly different from other northern Patagonian Maastrichtian units (e.g., Allen Formation, Los Alamitos Formation), which are dominated by saltasaurine sauropods and abelisaurids, suggesting potential temporal or environmental distinctions.[3][4]

Geology

The Chorrillo Formation is embedded within the foreland Magallanes Basin, shaped by Late Cretaceous Andean orogeny driven by subduction along the Pacific margin. Tectonic subsidence, coupled with sediment supply from the rising Patagonian Andes, facilitated deposition.[4] Lithologically, the unit comprises fine- to coarse-grained clastics: mudstones and siltstones dominate floodplain facies, often organic-rich and pedogenically modified; sandstones, arkosic to lithic, form channel fills with cross-bedding; and conglomerates appear as basal lags.[5] Clay mineralogy is dominated by smectite, with minor illite and kaolinite, reflecting volcanic ash alteration. Sedimentologically, it records a meandering fluvial system on a distal floodplain, featuring fining-upward channel sequences, crevasse splays, and overbank deposits.[3] High-frequency paleosol stacking indicates local hydrologic and topographic variability, while low-frequency patterns suggest avulsion-driven progradation. Regional ties extend to the Dorotea Formation in Chile, reflecting basin-wide tectonic influences.[5]

Paleoenvironment

Modern equivalent, Valdivian Rainforest

Paleoenvironmental reconstructions indicate a temperate-warm, seasonally humid climate with mean annual temperatures of 10-18°C and precipitation of 900-1500 mm, inferred from paleosols, clay mineralogy, and biotic indicators.[6] Depositional settings encompass low-energy fluvial floodplains with paludal and lacustrine elements, marked by poor drainage, hydromorphic paleosols (e.g., vertisols, histosols), and redoximorphic features such as mottling and iron nodules, suggesting fluctuating water tables and episodic anoxia.[5] Aquatic communities feature Nymphaeaceae, Salviniaceae, Salviniaceae, and Zygnemataceae, indicative of freshwater ponds and swamps, while terrestrial elements include conifers, ferns, and angiosperms in riparian zones.[7] Floral phytozones, derived from palynological data, comprise five assemblages across the section, exhibiting quantitative variations in taxa proportions without major turnover: lower zones are fern-dominated (e.g., Cyatheaceae), transitioning to angiosperm-rich upper zones (e.g., Proteaceae, Nothofagidites), attributed to subtle humidity fluctuations or habitat gradients relative to fluvial channels.[8] These patterns align with Maastrichtian assemblages in southern Gondwanan basins (e.g., La Anita, Lefipán Formation), reflecting angiosperm diversification and Weddellian Province expansion.[3][8]

Fossils

Dinosaur remains are among the fossils that have been recovered from the formation.

Fossils recovered from the formation are:

Color key
Taxon Reclassified taxon Taxon falsely reported as present Dubious taxon or junior synonym Ichnotaxon Ootaxon Morphotaxon
Notes
Uncertain or tentative taxa are in small text; crossed out taxa are discredited.

Arthropoda

Decapoda

Genus Species Locality Material Notes Images
Astacidea[9] Indeterminate "Locality 4" MPM 21516, three gastroliths Previously identified as crushing fish teeth
Extant example

Insects

Genus Species Locality Material Notes Images
Coelolepida[10] Indeterminate La Anita farm MPM-Pal 21835-3:W34/1, MPM-Pal 21835-23:W55/0, MPM-Pal 21835-22:R52/2,MPM-Pal 21835-20:Z47/3. MPM-Pal 21835-22: G55/0; MPM-Pal 2183-21: E42/3 A Glossatan Lepidopteran
Diamesinae[10] "Morphotype 3" La Anita farm MPM-Pal 21835-9:E34/4, MPM-Pal 21835-19: V34/4 A Chironomidae Nematoceran. Similar to the living genus Paraheptagyia
Ephemeroptera[10] Indeterminate La Anita farm MPM-Pal 21835-9:W37/1 A Mayfly
Lepidoptera[10] Indeterminate La Anita farm MPM-Pal 21835-10:X30/2, MPM-Pal 21835-7:S27/2
Orthocladiinae[10] "Morphotype 1" La Anita farm MPM-Pal 21835-24: M31/0, MPM-Pal 21835-3: P46/4 A Chironomidae Nematoceran
"Morphotype 2" La Anita farm MPM-Pal 21835-25: V26/3
"Morphotype 5" La Anita farm MPM-Pal 21835-9: Q58/3 It resembles modern Cricotopus
Tanypodinae[10] "Morphotype 4" La Anita farm MPM-Pal 21835-2:Y40/3 A Chironomidae Nematoceran

Molluscs

Gastropods

Genus Species Locality Material Notes Images
Allopeas[11] A. agnolini "Locality 4" Shells An achatinid gastropod
Extant specimen
Bocourtia[11] B. leonardodavincii "Locality 4" Shells A bulimulid gastropod
Extant specimen
Coelostemma[11] C. patagonica "Locality 4" Shells A holospirid gastropod
Holospira[4] H. sp. "Locality 4" Shells An urocoptidae gastropod
Leptinaria[11] Cf.L. sp. "Locality 4" Shells An achatinid gastropod
Extant specimen
Lilloiconcha[11] L. novasi "Locality 4" Shells A charopid gastropod
Naesiotus[11] Cf.N. sp. "Locality 4" Shells A bulimulid gastropod
Extant specimen
Physa[4] P. sp. "Locality 4" Shells A physidae gastropod
Extant specimen
Physella[11] Cf.P. sp. "Locality 4" Shells A physidae gastropod
Extant specimen
Pleuroceridae[4] Indeterminate "Locality 4" Shells Incertade Sedis
Pomacea[4] P. sp. "Locality 4" Shells An ampullariidae gastropod
Extant specimen
Potamolithus[11] P. fabiani "Locality 4" Shells A tateid gastropod
Extant specimen
Stenophysa[4] S. sp. "Locality 4" Shells A physidae gastropod

Bivalves

Color key
Taxon Reclassified taxon Taxon falsely reported as present Dubious taxon or junior synonym Ichnotaxon Ootaxon Morphotaxon
Notes
Uncertain or tentative taxa are in small text; crossed out taxa are discredited.
Genus Species Locality Material Notes Images
Chlamys[4] “C.” salamanca "Locality 5" Shells A Pectinidae bivalve
Extant specimen
Cubitostrea[4] C. ameghinoi "Locality 5" Shells An Ostreidae bivalve
Extant specimen
Gryphaeostrea[4] G. callophyla "Locality 5" Shells A Gryphaeidae bivalve

Fishes

Genus Species Locality Material Notes Images
Lamniformes[4] Indeterminate "Locality 5" Isolated upper lateral tooth Similar to Cretalamna.[3]
Vidalamiinae[4] Indeterminate "Locality 2" Isolated tooth An amiid fish

Amphibians

Genus Species Locality Material Notes Images
Anura[4] Indeterminate "Locality 4" MPM 21518, isolated tibiofibula
Calyptocephalella[3][4] C. gayi "Locality 2" Incomplete left maxilla, incomplete right maxilla, incomplete right ilium, proximal right radioulna, left proximal end of tibiofibula and proximal half of urostyle
Extant specimen
Calyptocephalellidae[4] Indeterminate "Locality 4" MPM 21519, distal end of right humerus with eroded distal margin
Pipoidea[3][4] Indeterminate "Locality 2" Distal end of right humerus Resembles, yet is clearly different, from Xenopus.[3]

Plesiosaurs

Genus Species Locality Material Notes Images
Elasmosauridae[3] Indeterminate "Locality 5" Incomplete cervical centrum and the lateral half of a very fragmentary dorsal vertebra May belong to the Calafate Formation

Lepidosaurs

Color key
Taxon Reclassified taxon Taxon falsely reported as present Dubious taxon or junior synonym Ichnotaxon Ootaxon Morphotaxon
Notes
Uncertain or tentative taxa are in small text; crossed out taxa are discredited.

Mosasaurs

Genus Species Locality Material Notes Images
Mosasauridae[3][4] Indeterminate "Locality 5" Seven teeth Its presence suggest marine influences

Rhynchocephalian

Genus Species Locality Material Notes Images
Notosphenos[12] N. finisterre "Monotreme site" Incomplete right maxilla preserving two completeand one partial teeth. A sphenodontid rhynchocephalian

Snakes

Genus Species Locality Material Notes Images
Alethinophidia[4][12] Indeterminate "Locality 4" MPM 21522 small vertebra
Anilioidea[3] Indeterminate "Locality 2" MPM-PV-22847, possible mid-precloacal vertebra
Coniophiidae[12] Indeterminate "Monotreme site" Nearly complete mid-posterior precloacal vertebra; Incomplete posterior precloacal vertebra Shares several features with Coniophis.
Madtsoiidae[12] Indeterminate "Monotreme site"; "Green Valley site" Incomplete mid-posterior precloacal vertebralacking most of the neural arch; Poorly preserved precloacal vertebra preserving the verteb-ral centrum
Rionegrophis[4] R. madtsoioides "Locality 2" A partial vertebra. A madtsoiid snake

Turtles

Genus Species Locality Material Notes Images
Chelidae[4] Indeterminate "Locality 2" MPM 21520, two peripheral and one costal plates Largest plates suggest a maximum carapace length of 40 cm.
Hydromedusinae[3] Indeterminate "Locality 2, 4, 5" Left peripheral 3 or 4, bridge peripheral, fragment indet; bridge peripheral, right hypoplastron, buttress, plastral fragment and ?plastral fragment
Meiolaniformes[3] Indeterminate "Locality 4" Distal end of a left humerus

Archosaurs

Crocodylomorphs

Genus Species Locality Material Notes Images
Kostensuchus[13] K. atrox Estancia La Anita Skull, jaws, incomplete postcranial skeleton A large-bodied, broad-snouted peirosaurid

Ornithischians

Genus Species Locality Material Notes Images
Hadrosauridae[4][14] Indeterminate La Anita Farm Two incomplete caudal centra
Isasicursor[4] I. santacrucensis "Locality 5" Multiple specimens including teeth, several vertebrae, humerus, thigh bones, shoulder blades, and limbs An elasmarian ornithopod
Isasicursor skeletal
Ornithopoda[4][14] Indeterminate La Anita Farm Proximal end of a femur A small sized ornithopod
Parankylosauria[14] Indeterminate La Anita Farm Five fragmentary caudal vertebrae, incomplete ribs, two incomplete humeri, and nine osteoderms

Sauropods

Genus Species Locality Material Notes Images
Fusioolithus[4] F. sp. "Locality 2" Eggshells Fusioolithidae oogenus
Lithostrotia[15] Indeterminate Río Leona MACN 5205 (3 middle–posterior caudal centra)
Nullotitan[4] N. glaciaris Estancia Alta Vista; "Locality 1, 2, 4, 5" A partial skeleton consisting of a third cervical vertebra, tail vertebrae, a neck rib, ribs, a left shoulder blade, the ends of a right thigh bone, a right shin, a right calf bone, and a right ankle bone A colossosaurian titanosaur framless
Titanosauria[4] Indeterminate "Locality 2, 4" Seven isolated teeth[4]

Theropods

Genus Species Locality Material Notes Images
Enantiornithes[3] Indeterminate "Locality 2, 4" Distal fragment of a pedal phalanx; Two isolated ungual pedal phalanges
Maip macrothorax
Kookne[4] K. yeutensis "Locality 2" An incomplete right coracoid, lacking the sternal end and the proximal end damaged An ornithuran bird similar to Anseriformes
Maip[16] M. macrothorax La Anita Farm Axis, several dorsal and caudal vertebrae, cervical and dorsal ribs, gastralia, a left coracoid, fragmentary scapula, partial right pubis, and partial metatarsal The largest megaraptorid theropod of South America and possibly the largest member of this group
Megaraptoridae[4] Indeterminate La Anita Farm Isolated teeth[4]
Noasauridae[4] Indeterminate "Locality 4" Isolated right pedal phalanx IV-2?[4] Similar to Velocisaurus and Vespersaurus
Prismatoolithidae[4] Indeterminate "Locality 4" Eggshells Theropod eggshells
Unenlagiidae[4] Indeterminate "Locality 4" An isolated dorsal vertebra, pedal ungual of digit II and phalanx III-2[4] Similar to Neuquenraptor
Yatenavis[17] Y. ieujensis Estancia La Anita Distal right humerus An enantiornithine bird

Mammals

Genus Species Locality Material Notes Images
Magallanodon[18] M. baikashkenke[19] Río de Las Chinas Valley and La Anita Farm Tentative first upper molariform, anterior end of a left dentary bearing part of its gliriform incisor, and upper mesial incisor from the right side A ferugliotheriid gondwanathere
M. terrwerr[20] "Locality 2" MPM-PV-23088, isolated right lower molariform; MPM-PV-23089, isolated right lower molariform A ferugliotheriid gondwanathere
Mammalia[4] Indeterminate "Locality 4" Two isolated caudal vertebrae Characters shared with mammals having long and gracile tails.
Meridiolestida[20] Indeterminate "Monotreme site" MPM-PV-21707, two fragments of a right dentary Small sized taxon similar to Reigitherium.
Mesungulatoidea[20] Indeterminate La Anita Farm MPM-PV-21708, fragment of left dentary with five alveoli corresponding with three teeth. Larger than any known Mesozoic meridiolestidan.[20]
Orretherium[20] O. tzei "Monotreme site" MPM-PV-23514, left upper second molariform. A member of Meridiolestida belonging to the family Mesungulatidae.
Patagorhynchus[21] P. pascuali "Monotreme site" Lower right molar attached to a fragment of the dentary A ornithorhynchid monotreme
Patagorhynchus holotype
Patagomaia[20][22] P. chainko La Anita Farm Partial left acetabulum and ischium; MPM-PV-23515, one fragment of thoracic vertebra, posterior half of one caudal vertebra and 38 indeterminate bones. A large therian
Patagomaia holotype

Flora

Color key
Taxon Reclassified taxon Taxon falsely reported as present Dubious taxon or junior synonym Ichnotaxon Ootaxon Morphotaxon
Notes
Uncertain or tentative taxa are in small text; crossed out taxa are discredited.
Algae
Genus Species Affinity / Family Image
Botryococcus[6][8] B. sp. Botryococcaceae
Cymatiosphaera[6][8] C. sp. Zygnemataceae
Leiosphaeridia[6][8] L. sp. Zygnemataceae
Ovoidites[6][8] O. parvus Zygnemataceae
O. spriggii Zygnemataceae
Schizosporis[6][8] S. reticulatus Zygnemataceae (aff. Spirogyra)
Sphaeroplea[8] S. striatocristata Sphaeropleaceae

Liverworts

Genus Species Affinity / Family Image
Aequitriradites[6][8] A. spinulosus Marchantiidae
Stoverisporites[6][8] S. microverrucatus Marchantiidae
Taurocusporites[6][8] T. segmentatus Marchantiidae
Triporoletes[6][8] T. reticulatus Marchantiidae
T. sp. Marchantiidae

Hornworts

Genus Species Affinity / Family Image
Foraminisporis[6][8] F. wonthaggiensis Anthocerotidae
F. sp. Anthocerotidae

Lycopods

Genus Species Affinity / Family Image
Camarozonosporites[6][8] C. ambigens Selaginellales
Ceratosporites[6][8] C. equalis Selaginellales
Densoisporites[6][8] D. velatus Selaginellales
Foveosporites[6][8] F. sp. Selaginellales
Retitriletes[6][8] R. austroclavatidites Lycopodiales
R. sp. cf. R. douglasii Lycopodiales

Ferns

Genus Species Affinity / Family Image
Azolla[6][8] A. cf. andreisii Salviniaceae
A. sp. Salviniaceae
Baculatisporites[6][8] B. comaumensis Osmundaceae
B. kachaikensis Osmundaceae
B. turbioensis Osmundaceae
cf. Cladophlebis[7] C. sp. Polypodiopsida
Clavifera[6][8] C. sp. Gleicheniaceae
Cyatheacidites[6][8] C. annulatus Dicksoniaceae (aff. Lophosoria)
Cyathidites[6][8] C. australis Cyatheaceae
C. minor Cyatheaceae
C. rafaeli Cyatheaceae
Dictyophyllidites[6][8] D. sp. cf. D. concavus Dipteridaceae / Matoniaceae
Gleicheniidites[6][8] G. sp. Gleicheniaceae
Leptolepidites[6][8] L. verrucatus Blechnaceae
Ornamentifera[6][8] O. sp. Gleicheniaceae
Osmundacidites[6][8] O. diazii Osmundaceae
Peromonolites[6][8] P. vellosus Blechnaceae

Non-conifer Seed Plants

Genus Species Affinity / Family Image
Cycadopites[6][8] C. follicularis Cycadales, Bennettitales, or Ginkgoales
Ephedripites[6][8] E. ambiguous Ephedrales
Equisetosporites[6][8] E. notensis Ephedrales
E. sp. cf. E. dudarensis Ephedrales
Vitreisporites[6][8] V. signatus Pteridosperms

Conifers

Genus Species Affinity / Family Image
Araucariacites[6][8] A. australis Araucariaceae
Callialasporites[6][8] C. turbatus Araucariaceae
Classopollis[6][8] C. sp. Cheirolepidiaceae
Cyclusphaera[6][8] C. psilata Araucariaceae
Dacrycarpites[6][8] D. australiensis Podocarpaceae
Dilwynites[6][8] D. tuberculatus Araucariaceae
Gamerroites[6][8] G. psilasaccus Podocarpaceae
Lygistepollenites[6][8] L. florinii Podocarpaceae
Microcachryidites[6][8] M. antarcticus Podocarpaceae (aff. Microcachrys)
Microcachrys
Peripollenites[6][8] P. sp. Cupressaceae
Phyllocladidites[6][8] P. mawsonii Podocarpaceae (aff. Lagarostrobos franklinii)
Lagarostrobos
Podocarpidites[6][8] P. elegans Podocarpaceae
P. elipticus Podocarpaceae
P. exiguus Podocarpaceae
P. herbstii Podocarpaceae
P. major Podocarpaceae
P. marwikii Podocarpaceae
P. otagoensis Podocarpaceae
P. sp. cf. P. langii Podocarpaceae
P. sp. cf. P. microreticuloidata Podocarpaceae
P. torquatus Podocarpaceae
P. verrucosus Podocarpaceae
Podocarpoxylon[4] P. dusenii Podocarpaceae
Trichotomosulcites[6][8] T. subgranulatus Podocarpaceae

Magnoliids

Genus Species Affinity / Family Image
Clavatimonocolpites[6][8] C. sp. Magnoliids

Monocots

Genus Species Affinity / Family Image
Arecipites[6][8] A. minutiscabratus Arecaceae
A. sp. cf. A. subverrucatus Arecaceae
Aff.Eolirion[7] "Morphotype 5" Palmae
Longapertites[6][8] L. proxapertitoides Arecaceae
Liliacidites[6][8] L. lanceolatus Liliales (aff. Lilium)
Lilium
L. variegatus Liliales (aff. Lilium)
L. vermireticulatus Liliales
L. sp. cf. L. kaitangataensis Liliaceae/Amaryllidaceae
Monocotyledonean[7] "Morphotype 6" Monocots
Nuphar[7] N.? sp. 1 Nymphaeaceae
Extant specimen
N.? sp. 2 Nymphaeaceae
Pandaniidites[6][8] P. typicus Araceae
Proxapertites[6][8] P. operculatus Araceae
Spinozonocolpites[6][8] S. hialinus Arecaceae

Eudicots

Genus Species Affinity / Family Image
Beauprea[6][8] B. elegansiformis Proteales
Beauprea drawing
Aff. Brachychiton[7] "Morphotype 4" Malvaceae
Extant specimen
Buttinia[6][8] B. andreevii Buxales
Clavamonocolpites[6][8] C. sp. ?Chloranthaceae
Coccoloba?[7] "Morphotype 1" Polygonaceae
Extant specimen
Constantinisporis[6][8] C. jacquei ?Chloranthaceae
Dicotetradites[6][8] D. sp. Ericales
Diporites[6][8] D. sp. ?Chloranthaceae
Doyleipollenites[6][8] D. sp. ?Chloranthaceae
Ericipites[6][8] E. scabratus Ericales
Eudicots[7] "Morphotype 3" Eudicots
Myrtaceidites[6][8] M. sp. cf. M. mesonesus Myrtales
Nothofagidites[6][8] N. saraensis Nothofagaceae (cf. Nothofagus)
Nothofagus
Nyssapollenites[6][8] N. sp. ?Chloranthaceae
Peninsulapollis[6][8] P. gillii Proteaceae (cf. Beauprea?)
P. truswelliae Proteaceae (cf. Beauprea?)
Polycolpites[6][8] P. sp. Pedaliaceae (?cf. Josephinia)
Proteacidites[6][8] P. parvus Proteaceae (cf. Bellendena montana)
Bellendena
P. subscabratus Proteaceae
P. sp. cf. P. tenuiexinus Proteaceae
P. sp. Proteaceae
Psilatricolporites[6][8] P. sp. cf. P. complanatius Tetrameristaceae
P. sp. Tetrameristaceae
Retidiporites[6][8] R. camachoi Proteaceae
Rhoipites[6][8] R. baculatus Ericales
R. minusculus Ericales
R. sp. cf. R. alveolatus Ericales
R. sp. 1 Ericales
R. sp. 2 Ericales
Rousea[6][8] R. patagonica Ericales
R. sp. cf. R. minuscula Ericales
R. sp. Ericales
Syncolporites[6][8] S. sp. Ericales
Tetracolpites[6][8] T. sp. ?Chloranthaceae
Triatriopollenites[6][8] T. lateflexus Proteaceae
Tricesticillus[6][8] T. americanus ?Chloranthaceae
Tricolpites[6][8] T. reticulatus Gunneraceae (Gunnera macrophylla)
Gunnera
T. sp. cf. T. brevicolpus Gunneraceae
T. sp. cf. T. inargutus Gunneraceae
T. sp. cf. T. lumbrerensis Gunneraceae
T. sp. 1 Ranunculaceae (cf. Clematis)
Clematis
Tricolporites[6][8] T. sp. 2 Ericales
T. sp. 3 Ericales
Tricolporoidites[6][8] T. sp. cf. T. eximius Ericales
Triorites[6][8] T. sp. cf. T. minusculus ?Chloranthaceae
Tubulifloridites[6][8] T. lilliei Ericales
Volkheimerites[6][8] V. labyrinthus ?Chloranthaceae

See also

References

  1. ^ Río Leona at Fossilworks.org
  2. ^ Marenssi et al., 2003, p.406
  3. ^ a b c d e f g h i j k l m n Moyano-Paz, D.; Rozadilla, S.; Agnolín, F.; Vera, E.; Coronel, M. D.; Varela, A. N.; Gómez-Dacal, A. R.; Aranciaga-Rolando, A. M.; D'Angelo, J.; Pérez-Loinaze, V.; Richiano, S.; Chimento, N.; Motta, M. J.; Sterli, J.; Manabe, M. (2022-02-01). "The uppermost Cretaceous continental deposits at the southern end of Patagonia, the Chorrillo Formation case study (Austral-Magallanes Basin): Sedimentology, fossil content and regional implications". Cretaceous Research. 130: 105059. doi:10.1016/j.cretres.2021.105059. ISSN 0195-6671.{{cite journal}}: CS1 maint: article number as page number (link)
  4. ^ a b c d e f g h i j k l m n o p q r s t u v w x y z aa ab ac ad ae af ag ah ai aj ak al Fernando Novas; Federico Agnolin; Sebastián Rozadilla; Alexis Aranciaga-Rolando; Federico Brissón-Eli; Matias Motta; Mauricio Cerroni; Martín Ezcurra; Agustin Martinelli; Julia D'Angelo; Gerardo Álvarez-Herrera; Adriel Gentil; Sergio Bogan; Nicolas Chimento; Jordi García-Marsà; Gastón Lo Coco; Sergio Miquel; Fatima Brito; Ezequiel Vera; Valeria Loinaze; Mariela Fernandez; Leonardo Salgado (2019). "Paleontological discoveries in the Chorrillo Formation (upper Campanian-lower Maastrichtian, Upper Cretaceous), Santa Cruz Province, Patagonia, Argentina". Revista del Museo Argentino de Ciencias Naturales. Nueva Series. 21 (2): 217–293. doi:10.22179/revmacn.21.655. hdl:11336/122229.
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